Tortoises are personal pets that have gained immense popularity around the world in recent years. And as evidenced by the proliferation of books and websites catering to the subject of captive tortoise maintenance, keepers of tortoises are constantly seeking information about providing their pets with optimal conditions. Many species adapt well to captivity, but pet tortoises are very much wild animals. Knowledge about their wild counterparts, therefore, may help tortoise keepers go a long way in satisfying the needs of their pet tortoises. The purpose of this article is to discuss aspects of the biology of wild tortoises – including evolutionary history, life history traits, physiological ecology, and social behaviors and communication – that can help keepers of tortoises provide a better existence for both captive and wild tortoises. In my third post, I discuss physiological ecology of tortoises.